· Professor, Iowa State University2012–
· Associate Professor, Iowa State University2007–2012
· Assistant Professor, UMass Medical School2001–2007
· Postdoctoral Research Associate, RNA Biology and Molecular Genetics, The University of Texas at Austin1998–2001
· Postdoctoral Research Associate, Biochemistry and Molecular Biology, Oregon State University1995–1998
· Postdoctoral Research Associate, Biochemistry, University Texas MD Anderson Cancer Center1993–1995
· Doctor of Philosophy (Ph.D.), Biochemistry, Russian Academy of Sciences1988–1993
· Master's Degree, Biochemistry, Banaras Hindu University1982–1985
· Bachelor's Degree, Chemistry (Honors), Banaras Hindu University1978–1983