· Associate Professor, University of Pennsylvania2011–
· Assistant Professor, University of Pennsylvania2003–2011
· Associate Reseach Scientist, Yale University School of Medicine2001–2003
· Postdoctoral Fellow, Yale University School of Medicine1996–2001
· Doctor of Philosophy (PhD), Inflammation, King's College London, U. of London1992–1996
· Master of Science (MSc), Cell Biology, The University of Manchester1989–1992
· BSc, Immunology, The University of Glasgow1984–1988