· Professor of the Graduate School, University of California, Berkeley2004–
· Professor of Integrative Biology, University of California, Berkeley1989–2003
· Professor of Zoology, University of California, Berkeley1980–1989
· Associate Professor of Biology and Zoology, University of California, Berkeley1976–1980
· Associate Dean, College of Letters and Science, University of California, Berkeley1975–1978
· Assistant Professor of Biology and Zoology, University of California, Berkeley1973–1976
· Assistant Dean, College of Letters and Science, University of California, Berkeley1972–1973
· Lecturer in Biology, University of California, Berkeley1969–1973
· Assistant Professor, University of Illinois, Chicago Circle1968–1969
· PhD Student, University of Southern California1968–
· Instructor, University of Illinois, Chicago Circle1966–1968
· MS in Computer Science, University of Southern California1964–
· Bachelor's degree, University of Southern California1961–