· Research Director, Institute of Biotechnology, University of Helsinki2020–
· Professor of Biotechnology, University of Helsinki2014–2018
· Academy Professor, University of Helsinki2009–2013
· Director, Biocenter Finland2008–2009
· Director and Professor, Institute of Biotechnology, University of Helsinki1990–2008
· Doctor of Biology, Tartu University1987–
· Professor (part-time), Tallinn Technical University1986–2016
· Head of the Department of Molecular Genetics, Estonian Academy of Sciences1980–1990
· Head of the Laboratory of Molecular Genetics, Estonian Academy of Sciences1977–1980
· PhD in Biology, Tartu University1975–
· MSc in Biochemistry and Molecular Biology, Tartu University1972–
· Research assistant & junior researcher, Tartu University1971–1977